1. The Western powers, particularly Great Britain, France, Russia, Germany, the United States, and Japan. These countries gained access to China's markets, resources, and labor force, and were able to exert significant political and economic influence over the country.
2. Chinese compradors, or merchants who collaborated with foreign businesses and governments. They acted as middlemen between foreign companies and Chinese merchants, and often made substantial profits from these transactions.
3. The Qing dynasty, which was able to use the unequal treaties to gain leverage in its negotiations with foreign powers. By granting concessions to foreign countries, the Qing dynasty was able to maintain its control over China and prevent further encroachment on its sovereignty.
4. The Chinese people, who benefited from increased trade, investment, and technology transfer from the foreign powers. However, these benefits were often limited to the coastal areas of China, and the vast majority of the Chinese population remained in poverty.