Rainforests aren't universally clay-rich. Sandy soils appear in specific geomorphic settings—and they reshape forest ecology.
Drainage: Rapid infiltration → no waterlogging. Nutrients: Low cation exchange capacity (CEC) → oligotrophic. Floristic distinctness: Specialized flora (sclerophyllous leaves, mycorrhizal dependence, carnivory). Fire: Dry-season flammability higher than clay forests.
Rio Negro basin (campinarana): white sand, stunted canopy (15-20m), high endemism (birds: Guianan cock-of-the-rock; plants: Brocchinia bromeliads). Upper Xingu: sandy soils underpin ancient anthropogenic dark earths (terra preta) — human-made fertility.
Borneo's kerangas (heath forests) on coastal sands: pitcher plants (Nepenthes), ant-plants (Myrmecodia), low stature. Threatened by oil palm on same soils.
Sandy patches are biodiversity hotspots within hotspots—refugia for specialists, corridors for generalists. Conservation planning must map soil heterogeneity, not just canopy cover.