Yes, Papua New Guinea (PNG) is unequivocally part of Australasia. This regional designation encompasses Australia, New Zealand, New Guinea, and the neighboring islands of the southwestern Pacific—a definition grounded in both tectonic geography and biogeography.
Australasia sits on the Indo-Australian Plate. New Guinea forms the northern edge of the Australian continental shelf, separated from Cape York Peninsula by the shallow Torres Strait (just 150 km wide, with depths under 50 m). During Pleistocene glacial maxima, sea levels dropped 120 m, exposing a land bridge—Sahul—that united Australia and New Guinea into a single continent. They share a continental crust, distinct from the oceanic plates of Melanesia to the east.
Alfred Russel Wallace identified the faunal boundary between Asia and Australasia running through the Lombok Strait (between Bali and Lombok) and continuing north between Borneo and Sulawesi. New Guinea falls squarely on the Australasian side. Marsupials (tree kangaroos, cuscus), monotremes (long-beaked echidnas), and bird families (birds of paradise, cassowaries) dominate—lineages with Australian affinities, not Asian. The flora similarly shows Gondwanan heritage: southern beeches (Nothofagus), conifers (Araucaria, Podocarpus), and diverse orchids.
While geographically Australasian, PNG is politically independent (since 1975) and culturally Melanesian. Its 800+ languages belong to the Austronesian and Papuan families, distinct from Aboriginal Australian or Polynesian groups. The term "Australasia" in geopolitical contexts sometimes excludes PNG, favoring "Australia and New Zealand" or "Oceania" (which includes Melanesia, Micronesia, Polynesia). But in physical geography, earth science, and ecology, PNG is Australasian.
Conservation planning, climate modeling, and evolutionary biology all treat the Australasian realm as a unit. The New Guinea highlands—rising to 4,509 m at Puncak Jaya—create climatic zones from tropical lowlands to alpine grasslands, hosting 5–10% of Earth's species on 0.5% of its land area. Understanding PNG as Australasian explains its unique biodiversity: it's an uplifted fragment of the Australian continent, not a volcanic island arc.